个人简介
姓名：毛娟      职称：教授    职务：副院长
联系电话：0931-7632399
电子邮箱：maojuan@gsau.edu.cn
邮寄地址：甘肃省兰州市安宁区营门村1号
办公地址：甘肃农业大学农科楼6楼613室
所在部门：园艺学院果树科学系

教育背景
2010.09-2014.06 甘肃农业大学，农学院，博士
2004.09-2007.06 甘肃农业大学，农学院，硕士
2000.09-2004.06 甘肃农业大学，园艺系，本科
工作履历
2020.12-至今 甘肃农业大学园艺学院，教授
2015.12-2020.11 甘肃农业大学园艺学院，副教授
2014.01-2015.11 甘肃农业大学园艺学院，讲师
2010.12 -2013.12 甘肃农业大学农学院，讲师
2007.07-2010.11 甘肃农业大学农学院，助教
学术兼职
1. 甘肃省园艺学会秘书长
2. 甘肃省现代水果产业体系专家
3. 甘肃省果树果品标准委员会副秘书长
教学工作
1. 研究生：果树种质创新与应用、园艺植物种质资源与利用
2. 研究专业选修课：高级果树育种学
研究领域
1. 果树生物技术与种质创新

获奖及荣誉

1. 河西走廊酿酒葡萄全产业链关键技术研发与应用，获甘肃省科技进步二等奖，1/14，2024年；

2. 甘肃省第二层次领军人次，2023年；

3. 甘肃省飞天学者特聘教授，2022年；

4. 甘肃优势林果种质创新及高质高效关键技术研发与示范应用，获甘肃省科技进步一等奖，5/15，2022年；

5. 第十届甘肃青年科技奖，2021年；

6. 甘肃省青年教师成才奖，2018年；

7. 葡萄抗寒砧木筛选与嫁接苗抗寒栽培技术集成示，获得甘肃省科技进步二等奖，2/12，2018 年；

8. 黄土高原雨养农业区苹果最佳建园模式研究，获得甘肃省科技进步二等奖，3/12，2016 年。

科研项目

1. ‘VabHLH71-VaGH3.6-like/VaLNK2’ 参与葡萄低温胁迫响应的分子机制，国家自然科学基金，2025年，主持，在研；

2. 智慧果园建设与高质高效生产模式应用，中央引导地方科技发展资金，2025，主持，在研；

3. 高等学校产业支撑计划项目-陇东黄土高原果畜草一体化循环模式建立与示范应用，甘肃省教育厅，2023年，主持，在研；

4. 丝路寒旱农业科技项目-苹果果园清洁生产与种养循环利用关键技术研发与示范，甘肃省农业农村厅，2023年，主持，在研；

5. 国家重点研发计划子课题-静宁现代矮砧苹果提质增效关键技术集成与示范，中华人民共和国科学技术部，主持，在研；

6. 葡萄JA信号介导的抗寒机理与抗寒品种的选育，甘肃省陇原创新青年，主持，在研；

7. 葡萄低温相关基因BAM3的克隆与功能鉴定, 甘肃农业大学“伏羲杰出人才培育计划”，主持，在研；

8. 葡萄赤霉素氧化酶基因家族的功能解析与抗低温机制研究，甘肃省自然基金重点项目，2022年，主持人，结题。

代表性成果
（一）期刊论文

1. Yongqing Feng, Min Li, Shixiong Lu, Miao Shao, Guoping Liang, Juan Mao*. Identification of the grape MAPKKK gene family and functional analysis of the VaMAPKKK15 gene under low temperature stress[J]. Plant Physiology and Biochemistry, 2025, 220: 109533.

2. Han Wang, Juanjuan Huang, Congcong Zhang, Yanmei Li, Huimin Gou, Guoping Liang, Zonghuan Ma, Juan Mao, Baihong Chen. Glucose enhanced lignin accumulation in grapevine stems via promoting phenylpropanoid biosynthesis[J]. Chemical and Biological Technologies in Agriculture, 2024, 11(1), 152.

3. Huimin Gou, Shixiong Lu, Guojie Nai, Weifeng Ma, Jiaxuan Ren, Lili Guo, Baihong Chen, Juan Mao*. The role of gibberellin synthase gene VvGA2ox7 acts as a positive regulator to salt stress in Arabidopsis thaliana[J]. BMC Plant Biology, 2024, 24(1):1051.

4. Huimin Gou, Shixiong , Lili Gu, Lili Ch, Min Li, Baozhen Zeng, Juanbo Yang, Baihong Chen, Juan Mao*. Evolution of PIN gene family between monocotyledons and dicotyledons and VvPIN1 negatively regulates freezing tolerance in transgenic Arabidopsis[J]. Physiologia Plantarum, 2024, 176(4):e14464.

5. Shixiong Lu, Xueting Zhou, Xu Huang, Baozhen Zeng, Huimin Gou, Weifeng Ma, Zonghuan Ma, Juan Mao, Baihong Chen*. Comparative analysis of the NF-Y transcription factor family identifies VaNF-YA6 as a positive regulator of salt and drought tolerance in grapevine[J]. Plant Stress, 2024, 14 , 100658.

6. Shixiong Lu, Min Li, Yongjuan Cheng, Huimin Gou, Lili Che, Guoping Liang, Juan Mao*. Genome-wide identification of Aux/IAA gene family members in grape and functional analysis of VaIAA3 in response to cold stress[J]. Plant Cell Reports, 2024, 43(11):265.

7. Yanmei Li, Guoping Liang, Shixiong Lu, Han Wang, Fanwei Zeng, Guojie Nai, Juan Mao, Baihong Chen*. Overexpression of VaSS4 negatively regulates cold tolerance by disturbing ROS balance and decreasing soluble sugar content[J].Scientia Horticulturae, 2024, 337, 113590.

8. Weifeng Ma, Lijuan Ma, Zonghuan Ma, Wenfang Li, Shixiong Lu, Huimin Gou, Juan Mao, Baihong Chen*. Profiling the lncRNA-miRNA-mRNA interaction network in the cold-resistant exercise period of grape (Vitis amurensis Rupr)[J].Chemical and Biological Technologies in Agriculture, 2024, 11(1), 143.

9. Fanwei Zeng, Yongqing Feng, Tian Wang, Xiyuan Ma, Shuzhen Jiao, Shangwen Yang, Miao Shao, Zonghuan Ma, Juan Mao, Baihong Chen*. The asymmetric expression of plasma membrane H+-ATPase family genes in response to pulvinus-driven leaf phototropism movement in Vitis vinifera[J]. Physiologia plantarum, 2024, 176(3):e14380.

10. BaoZhen Zeng, XueTing Zhou, HuiMin Gou, LiLi Che, ShiXiong Lu, JuanBo Yang, YongJuan Cheng, GuoPing Liang, Juan Mao*. Molecular Evolution of SNAREs in Vitis vinifera and Expression Analysis under Phytohormones and Abiotic Stress[J]. International Journal of Molecular Sciences, 2024, 25(11):5984.

11. Miao Shao, Yongqing Feng, Shangwen Yang, Tong Feng, Fanwei Zeng, Shixiong Lu, Zonghuan Ma, Baihong Chen, Juan Mao*. Molecular evolution of Phytocyanin gene and analysis of expression at different coloring periods in apple (Malus domestica)[J]. International Journal of Molecular Sciences, 2024, 24(1):374.

12. Weifeng Ma, Shixiong Lu, Wenfang Li, Guojie Nai, Zonghuan Ma, Yanmei Li, Baihong Chen, Juan Mao*. Transcriptome and metabolites analysis of water‐stressed grape berries at different growth stages[J]. Physiologia Plantarum, 2023, 175(3), e13910.

13.Yanmei Li, Guoping Liang, Guojie Nai, Shixiong Lu, Weifeng Ma, Zonghuan Ma, Juan Mao, Baihong Chen*. VaSUS2 confers cold tolerance in transgenic tomato and Arabidopsis by regulation of sucrose metabolism and ROS homeostasis[J]. Plant Cell Reports, 2023, 42(3):505-520.

14. Miao Shao, Ping Wang, Huimin Gou, Ma Zonghuan, Chen Baihong, Juan Mao. Identification and expression analysis of the SKP1-like gene family under phytohormone and abiotic stresses in apple (Malus domestica) [J]. International Journal of Molecular Sciences, 2023, 24(22), 16414.

15. Lili Guo, Shixiong Lu, Tao Liu, Guojie Nai, Jiaxuan Ren, Huimin Gou, Baihong Chen, Juan Mao*. Genome-wide identification and abiotic stress response analysis of PP2C gene family in woodland and pineapple strawberries[J]. International Journal of Molecular Sciences, 2023, 24: 4049.

16. Jiaxuan Ren, Pan Zhang, Yingbao Dai, Xiaohuan Liu, Shixiong Lu, Lili Guo, Huimin Gou Juan Mao*. Evolution of the 14–3–3 gene family in monocotyledons and dicotyledons and validation of MdGRF13 function in transgenic Arabidopsis thaliana[J]. Plant Cell Reports, 2023, 1-20.

17. Jiaxuan Ren, Li Feng, Lili Guo, Huimin Gou, Shixiong Lu, Juan Mao*. Genome-wide identification and expression analysis of the BURP domain-containing genes in Malus domestica[J]. Physiology and Molecular Biology of Plants, 2023, 1-15.

18. Jiaxuan Ren, Wenfang Li, Zhigang Guo, Zonghuan Ma, Dongshi Wan, Shixiong Lu, Lili Guo, Huimin Gou, Baihong Chen, Juan Mao*. Whole-genome resequencing and transcriptome analyses of four generation mutants to reveal spur-type and skin-color related genes in apple (Malus domestica Borkh. Cv. Red delicious)[J]. BMC Plant Biology, 2023, 23(1), 607.

19. Guoping Liang, Yingjun Hou, Han Wang, Ping Wang, Juan Mao*, Baihong Chen*. VaBAM1 weakens cold tolerance by interacting with the negative regulator VaSR1 to suppress β-amylase expression[J]. International Journal of Biological Macromolecules, 2023, 15;225:1394-1404.

20. Yanmei Li, Guoping Liang, Guojie Nai, Shixiong Lu, Weifeng Ma, Zonghuan Ma , Juan Mao*, Baihong Chen*. VaSUS2 confers cold tolerance in transgenic tomato and Arabidopsis by regulation of sucrose metabolism and ROS homeostasis[J]. Plant Cell Reports, 2023, doi: 10.1007/s00299-022-02972-w.

21. Ping Wang, Shixiong Lu, Xuejing Cao, Zonghuan Ma, Baihong Chen*, Juan Mao*. Physiological and transcriptome analyses of the effects of excessive water deficit on malic acid accumulation in apple. Tree Physiology, 2022, 00: 1-16.

22. Shixiong Lu, Honghong He, Ping Wang, Huiming Gou, Xuejing Cao, Zonghuan Ma, Baihong Chen, Juan Mao*. Evolutionary relationship analysis of STARD gene family and VvSTARD5 improves tolerance of salt stress in transgenic tomatoes[J]. Physiologia Plantarum. 2022, 174: e13772.

23. Guoping Liang, Zonghuan Ma, Shixiong Lu, Weifeng Ma, Lidan Feng, Juan Mao*, Baihong Chen*. Temperature‑phase transcriptomics reveals that hormones and sugars in the phloem of grape participate in tolerance during cold acclimation[J]. Plant Cell Reports (2022) 41:1357–1373.

24. Guojie Nai, Guoping Liang, Weifeng Ma, Shixiong Lu, Yanmei Li, Huimin Gou, Lili Guo, Baihong Chen, Juan Mao*. Overexpression VaPYL9 improves cold tolerance in tomato by regulating key genes in hormone signaling and antioxidant enzyme[J]. BMC Plant Biology, 2022, 22: 344.

25. Huimin Gou, Guojie Nai, Shixiong Lu, Weifeng Ma, Baihong Chen, Juan Mao*. Genome‑wide identification and expression analysis of PIN gene family under phytohormone and abiotic stresses in Vitis Vinifera L[J]. Physiol Mol Biol Plants, 2022, 28: 1905-1919.

26. Lili Che, Shixiong Lu, Guoping Liang, Huimin Gou, Min Li, Baihong Chen , Juan Mao*. Identification and expression analysis of the grape pentatricopeptide repeat (PPR) gene family in abiotic stress[J]. Physiol Mol Biol Plants, 28: 1849–1874 .

27. Shixiong Lu, Ping Wang, Guojie Nai, Yanmei Li, Yanli Su, Guoping Liang, Baihong Chen, Juan Mao*. Insight into VvGH3 genes evolutional relationship from monocotyledons and dicotyledons reveals that VvGH3-9 negatively regulates the drought tolerance in transgenic Arabidopsis[J]. Plant Physiology and Biochemistry, 2022, 172: 70-86.

28. Guoping Liang, Honghong He, Guojie Nai, Lidan Feng, Yanmei Li, Qi Zhou, Zonghuan Ma, Yuan Yue, Baihong Chen, Juan Mao*. Genome-wide identification of BAM genes in grapevine (Vitis vinifera L.) and ectopic expression of VvBAM1 modulating soluble sugar levels to improve low-temperature tolerance in tomato[J]. BMC Plant Biology, 2021, 21: 156.

29. Honghong He, Guoping Liang, Shixiong Lu, Pingping Wang, Tao Liu, Zonghuan Ma, Cunwu Zuo, Xiaomei Sun, Baihong Chen, Juan Mao*. Genome-wide identification and expression analysis of GA2ox, GA3ox, and GA20ox are related to Gibberellin Oxidase Genes in grape (Vitis vinifera L.) [J]. Genes, 2019, 10: 680.

30. Xin Zhao, Wenfang Li, Ying Wang, Zonghuan Ma, Shijin Yang, Qi Zhou, Juan Mao*, Baihong Chen*. Elevated CO2 concentration promotes photosynthesis of grape (Vitis vinifera L. cv. ‘Pinot noir’) plantlet in vitro by regulating RbcS and Rca revealed by proteomic and transcriptomic profiles[J]. BMC Plant Biology, 2019, 19: 42.

31. Wenfang Li, Juan Mao, Xinwen Li, Jing Su, Mohammed Mujitaba Dawuda, Zonghuan Ma, Cunwu Zuo, Zeshan An, Baihong Chen*. Effects of CEPA and 1-MCP on flower bud differentiation of apple cv. ‘Nagafu No. 2’grafted on different rootstocks[J]. Journal of Plant Growth Regulation, 2019, 38(3): 842-854.

32. Zonghuan Ma, Wenfang Li, Juan Mao, Wei Li 1, Cunwu Zuo, Xin Zhao, Mohammed Mujitaba Dawuda, Xingyun Shi, Baihong Chen*. Synthesis of light-inducible and light-independent anthocyanins regulated by specific genes in grape ‘Marselan’ (Vitis. vinifera L.)[J]. Peer J, 2019, 7: e6521.

33. Juan Mao, Wenfang Li, Baoqin Mi, Mohammed Mujitaba Dawuda, Zonghuan Ma, Yongmei Zhang, Baihong Chen*. Transcriptome analysis revealed glucose application affects plant hormone signal transduction pathway in "Red Globe" grape plantlets[J]. Plant Growth Regulation, 2018, 84: 45-56.

34. Jiangping Bai, Juan Mao, Hongyu Yang, Awais Khan, Aqi Fan, Siyan Liu, Junlian Zhang, Di Wang, Huijuan Gao and Jinlin Zhang. Sucrose non-ferment 1 related protein kinase 2 (SnRK2) genes could mediate the stress responses in potato (Solanum tuberosum L.)[J]. BMC Genetics, 2017, 18: 41.

35. Juan Mao, Wenfang Li, Baoqin Mi, Mohammed Mujitaba Dawuda, Alejandro Calderón-Urrea, Zonghuan Ma, Yongmei Zhang, Baihong Chen*. Different exogenous sugars affect the hormone signal pathway and sugar metabolism in “Red Globe” (Vitis vinifera L.) plantlets grown in vitro as shown by transcriptomic analysis[J]. Planta, 2017, 246: 537-552.

36. Jing Ren, Juan Mao, Cunwu Zuo, Alejandro Calderón-Urrea, Mohammed M.D., Xin Zhao, Xinwen Li, Baihong Chen*. Significant and unique changes in phosphorylation levels of four phosphoproteins in two apple rootstock genotypes under drought stress[J]. Molecular Genetic and Genomics, 2017, 292:1307-1322.

37. Baihong Chen, Juan Mao, Baona Huang, Baoqin Mi, Yulian Liu, Zijing Hu and Zonghuan Ma. Effect of bagging and time of harvest on fruit quality of 'Red Fuji' apple in high altitude area in China[J]. Fruits, 2017, 72 (1): 36-46.

38. Jing Ren, Xinwen Li, Juan Mao, Cunwu Zuo, Xin Zhao, Baihong *. Physiological and quantitative phosphoproteome analyses of drought stress-induced mechanisms in Malus baccata (L.) Borkh[J]. Biochemical Systematics and Ecology, 2017, 72: 47-55.

39. Yulian Liu, Nianlai Chen, Zonghuan Ma, Fei Che, Juan Mao, Baihong Chen*. The Changes in color, soluble sugars, organic acids, anthocyanins and aroma components in “Starkrimson” during the ripening period in china[J]. Molecules, 2016, 21(6): 812.

40. 史光玲, 董媛媛, 李诗美, 曾宝珍，卢世雄，梁国平，陈佰鸿，毛娟*. 叶面肥对黄土高原苹果‘烟富3号’果实品质的影响[J]. 干旱地区农业研究, 2025, 43(01): 103-116.

41. 杨娟博, 郭丽丽, 卢世雄, 苟惠敏,王 帅珽, 曾宝珍,毛娟*. 草莓FaGH3.17基因的克隆及功能分析[J]. 园艺学报, 2024, 51(11): 2483-2494.

42.梁国平, 曾宝珍, 刘铭, 边志远, 陈佰鸿, 毛娟*. 山葡萄VaSR基因家族的鉴定及VaSR1抗寒功能验证与互作蛋白筛选[J]. 园艺学报, 2025, 52(01): 37-50.

43.马宗桓, 李玉梅, 韦霞霞, 李文芳, 陈佰鸿, 毛娟*. 河西走廊不同产地‘美乐’葡萄品质及香气物质的差异分析[J]. 园艺学报, 2024,51(05): 1083-1098.

44.曾宝珍, 成永娟, 车莉莉, 杨娟博, 卢世雄, 梁国平, 吴志国, 赵毅, 毛娟*. 纳米零价铁对武威产区黑比诺葡萄新梢和叶片生长及光合特性的影响[J]. 果树学报, 2024, 41(03): 481-493.

45.董媛媛, 任家玄, 车莉莉, 郭丽丽, 成永娟, 毛娟*. 苹果CAMTA转录因子家族的鉴定与表达分析[J]. 植物生理学报, 2024, 60(01): 155-166.

46.刘芳, 梁国平, 王晗, 任家玄, 陈佰鸿, 毛娟*. 森林草莓SAUR基因家族的鉴定与表达分析[J]. 农业生物技术学报, 2024, 32(01): 91-106.

47.郭丽丽, 卢世雄, 乃国洁, 任家玄, 苟惠敏, 陈佰鸿, 毛娟*. 凤梨草莓和森林草莓GH3基因家族的鉴定与表达分析[J]. 植物生理学报, 2023, 59(03): 579-598.

48.李敏, 韩丽娜, 梁国平, 陈佰鸿, 毛娟*. 苹果MdMAP70-1克隆及异源过表达番茄的抗旱性分析[J]. 西北植物学报, 2023, 43(02): 181-189.

49.任家玄, 李艳梅, 马维峰, 吴宙, 毛娟*. 苹果14-3-3基因家族的鉴定与MdGRF13的功能分析[J]. 果树学报, 2023, 40(03): 405-421.

50. 郭丽丽, 卢世雄, 乃国洁, 马维峰, 毛娟*. 草莓PYL基因家族的鉴定与非生物胁迫下的表达分析[J]. 农业生物技术学报, 2023, 30(12):2315-2332.

51. 刘铭, 杨尚雯, 马维峰等. 葡萄蔗糖转化酶基因家族的生物信息学及响应逆境胁迫分析[J]. 西北植物学报, 2023, 43(11): 1801-1812.

52. 马维峰, 李艳梅, 马宗桓, 陈佰鸿, 毛娟*. 苹果POD家族基因的鉴定与 MdPOD15的功能分析[J]. 园艺学报, 2022, 49(6): 1181-1199.

53. 任家玄, 李艳梅, 马维峰, 吴宙, 毛娟*. 苹果14-3-3基因家族的鉴定与MdGRF13的功能分析[J]. 果树学报, 2022 .

（二）出版专著
1. 葡萄优质高效栽培技术，主编，甘肃科技出版社，2020；
2. 葡萄，主编，甘肃科技出版社，2018；
3. 植物组织培养，参编，中国林业出版社，2020；
4. 葡萄分子耕田，参编，中国林业出版社，2019；
5. 苹果标准化栽培技术，参编，甘肃科技出版社，2011。

（三）授权专利

1. 拉枝辅助垫片，ZL 2020 2 0042224X，第一完成人；
2. 可调式果树拉枝器，ZL 2020 2 0041554.7，第一完成人；
3. 拉枝精准省力开角器，ZL 2020 2 0041583.3，第一完成人；
4. 一种葡萄种植用培育架，ZL 2020 2 0045555.9，第一完成人；

（四）制定标准
1. 葡萄抗寒性评价规范 DB62/T4485-2021